September, 1939, and Britain is at war. Ruth Aspinall, a gifted pilot, is determined to join the Air Transport Auxiliary, an organization of civilian pilots who ferry aircraft to wherever they're needed. Meanwhile, over in America, Jack and Lucy Nelson, both experienced pilots themselves, are keen to join too. After a perilous journey by sea, Jack is soon having his first experience of the London Blitz before being posted to White Waltham, where he quite literally bumps into Ruth. By 1944 the tide of war is changing. Jack is sent to France to deliver a Spitfire, but his plane never arrives.